A million wildebeest… each one driven by the same ancient rhythm, fulfilling its instinctive role in the inescapable cycle of life: a frenzied three-week bout of territorial conquests and mating; survival of the fittest as 40km (25 mile) long columns plunge through crocodile-infested waters on the annual exodus north; replenishing the species in a brief population explosion that produces more than 8,000 calves daily before the 1,000 km (600 mile) pilgrimage begins again.

Tanzania’s oldest and most popular national park, the Serengeti is famed for its annual migration, when some six million hooves pound the open plains, as more than 200,000 zebra and 300,000 Thomson’s gazelle join the wildebeest’s trek for fresh grazing.

Yet even when the migration is quiet, the Serengeti offers arguably the most scintillating game-viewing in Africa: great herds of buffalo, smaller groups of elephant and giraffe, and thousands upon thousands of eland, topi, kongoni, impala and Grant’s gazelle.

The spectacle of predator versus prey dominates Tanzania’s greatest park. Golden-maned lion prides feast on the abundance of plain grazers. Solitary leopards haunt the acacia trees lining the Seronera River, while a high density of cheetahs prowls the southeastern plains.

Almost uniquely, all three African jackal species occur here, alongside the spotted hyena and a host of more elusive small predators,ranging from the insectivorous aardwolf to the beautiful serval cat.But there is more to Serengeti than large mammals. Gaudy agama lizards and rock hyraxes scuffle around the surfaces of the park’s isolated granite koppies.

A full 100 varieties of dung beetle have been recorded, as have 500-plus bird species, ranging from the outsized ostrich and bizarre secretary bird of the open grassland, to the black eagles that soar effortlessly above the Lobo Hills.

As enduring as the game-viewing is the liberating sense of space that characterises the Serengeti Plains, stretching across sunburnt savannah to a shimmering golden horizon at the end of the earth. Yet, after the rains, this golden expanse of grass is transformed into an endless green carpet flecked with wildflowers. And there are also wooded hills and towering termite mounds, rivers lined with fig trees and acacia woodland stained orange by dust.

Popular the Serengeti might be, but it remains so vast that you may be the only human audience when a pride of lions masterminds a siege, focussed unswervingly on its next meal.

About Serengeti
Size: 14,763 sq km (5,700 sq miles).
Location: 335km (208 miles) from Arusha, stretching north to Kenya and bordering Lake Victoria to the west.

What to do
Hot air balloon safaris, Maasai rock paintings and musical rocks.
Visit neighbouring Ngorongoro Crater, Olduvai Gorge, Ol Doinyo Lengai volcano and Lake Natron’s flamingos.

When to go
To follow the wildebeest migration, December-July. To see predators, June-October
Accommodation

NOTE
The route and timing of the wildebeest migration is unpredictable. Allow at least three days to be assured of seeing them on your visit – longer if you want to see the main predators as well.

Kilimanjaro. The name itself is a mystery wreathed in clouds. It might mean Mountain of Light, Mountain of Greatness or Mountain of Caravans. Or it might not. The local people, the Wachagga, don’t even have a name for the whole massif, only Kipoo (now known as Kibo) for the familiar snowy peak that stands imperious, overseer of the continent, the summit of Africa.

Kilimanjaro, by any name, is a metaphor for the compelling beauty of East Africa. When you see it, you understand why. Not only is this the highest peak on the African continent; it is also the tallest free-standing mountain in the world, rising in breathtaking isolation from the surrounding coastal scrubland – elevation around 900 metres – to an imperious 5,895 metres (19,336 feet).

Kilimanjaro is one of the world’s most accessible high summits, a beacon for visitors
from around the world. Most climbers reach the crater rim with little more than a walking stick, proper clothing and determination.

And those who reach Uhuru Point, the actual summit, or Gillman’s Point on the lip of the crater, will have earned their climbing certificates.

And their memories.

But there is so much more to Kili than her summit. The ascent of the slopes is a virtual climatic world tour, from the tropics to the Arctic.

Even before you cross the national park boundary (at the 2,700m contour), the cultivated footslopes give way to lush montane forest, inhabited by elusive elephant, leopard, buffalo, the endangered Abbot’s duiker, and other small antelope and primates. Higher still lies the moorland zone, where a cover of giant heather is studded with otherworldly giant lobelias.

Above 4,000m, a surreal alpine desert supports little life other than a few hardy mosses and lichen. Then, finally, the last vestigial vegetation gives way to a winter wonderland of ice and snow – and the magnificent beauty of the roof of the continent.

About Kilimanjaro National Park

Size: 755 sq km (292 sq miles).
Location: Northern Tanzania, near the town of Moshi.

What to do

Six usual trekking routes to the summit and other more-demanding mountaineering routes. Day or overnight hikes on the Shira plateau. Nature trails on the lower reaches.

Trout fishing.

Visit the beautiful Chala crater lake on the mountain’s southeastern slopes.

When to go

Clearest and warmest conditions from December to February, but also dry (and colder) from July-September.

Accommodation
Huts and campsites on the mountain.

Set deep in the heart of the African interior, inaccessible by road and only 100km (60 miles) south of where Stanley uttered that immortal greeting “Doctor Livingstone, I presume”, is a scene reminiscent of an Indian Ocean island beach idyll.

Silky white coves hem in the azure waters of Lake Tanganyika, overshadowed by a chain of wild, jungle-draped peaks towering almost 2km above the shore: the remote and mysterious Mahale Mountains.

Mahale Mountains, like its northerly neighbour Gombe Stream, is home to some of Africa’s last remaining wild chimpanzees: a population of roughly 800, habituated to human visitors by a Japanese research project founded in the 1960s.

Tracking the chimps of Mahale is a magical experience. The guide’s eyes pick out last night’s nests – shadowy clumps high in a gallery of trees crowding the sky. Scraps of half-eaten fruit and fresh dung become valuable clues, leading deeper into the forest. Butterflies flit in the dappled sunlight.

Then suddenly you are in their midst: preening each other’s glossy coats in concentrated huddles, squabbling noisily, or bounding into the trees to swing effortlessly between the vines.

The area is also known as Nkungwe, after the park’s largest mountain, held sacred by the local Tongwe people, and at 2,460 metres (8,069 ft) the highest of the six prominent points that make up the Mahale Range. And while chimpanzees are the star attraction, the slopes support a diverse forest fauna, including readily observed troops of red colobus, red-tailed and blue monkeys, and a kaleidoscopic array of colourful forest birds.

You can trace the Tongwe people’s ancient pilgrimage to the mountain spirits, hiking through the montane rainforest belt – home to an endemic race of Angola colobus monkey – to high grassy ridges chequered with alpine bamboo. Then bathe in the impossibly clear waters of the world’s longest, second-deepest and least-polluted freshwater lake – harbouring an estimated 1,000 fish species – before returning as you came, by boat.

About Mahale Mountains National Park
Size: 1,613 sq km (623 sq miles).
Location: Western Tanzania, bordering Lake Tanganyika.

What to do
Chimp tracking (allow two days); hiking; camping safaris; snorkelling; fish for your dinner.

When to go
Dry season (May-October) best for forest walks although no problem in the light rains of October/November.

NOTE:
The same rules for chimpanzee viewing at Gombe Stream apply at Mahale

Isolated, untrammelled and seldom visited, Katavi is a true wilderness, providing the few intrepid souls who make it there with a thrilling taste of Africa as it must have been a century ago. Tanzania’s third largest national park, it lies in the remote southwest of the country, within a truncated arm of the Rift Valley that terminates in the shallow, brooding expanse of Lake Rukwa.

The bulk of Katavi supports a hypnotically featureless cover of tangled brachystegia woodland, home to substantial but elusive populations of the localised eland, sable and roan antelopes. But the main focus for game viewing within the park is the Katuma River and associated floodplains such as the seasonal Lakes Katavi and Chada. During the rainy season, these lush, marshy lakes are a haven for myriad waterbirds, and they also support Tanzania’s densest concentrations of hippo and crocodile.

It is during the dry season, when the floodwaters retreat, that Katavi truly comes into its own. The Katuma, reduced to a shallow, muddy trickle, forms the only source of drinking water for miles around, and the flanking floodplains support game concentrations that defy belief. An estimated 4,000 elephants might converge on the area, together with several herds of 1,000-plus buffalo, while an abundance of giraffe, zebra, impala and reedbuck provide easy pickings for the numerous lion prides and spotted hyena clans whose territories converge on the floodplains.

Katavi’s most singular wildlife spectacle is provided by its hippos. Towards the end of the dry season, up to 200 individuals might flop together in any riverine pool of sufficient depth. And as more hippos gather in one place, so does male rivalry heat up – bloody territorial fights are an everyday occurrence, with the vanquished male forced to lurk hapless on the open plains until it gathers sufficient confidence to mount another challenge.

About Katavi National Park
Size: 4,471 sq km (1,727 sq miles).
Location: Southwest Tanzania, east of Lake Tanganyika.
The headquarters at Sitalike lie 40km (25 miles) south of Mpanda town.

What to do.

Walking, driving and camping safaris.
Near Lake Katavi, visit the tamarind tree inhabited by the spirit of the legendary hunter Katabi (for whom the park is named) – offerings are still left here by locals seeking the spirit’s blessing.

When to go
The dry season (May-October).
Roads within the park are often flooded during the rainy season but may be passable from mid-December to February

An excited whoop erupts from deep in the forest, boosted immediately by a dozen other voices, rising in volume and tempo and pitch to a frenzied shrieking crescendo. It is the famous ‘pant-hoot’ call: a bonding ritual that allows the participants to identify each other through their individual vocal stylisations.

To the human listener, walking through the ancient forests of Gombe Stream, this spine-chilling outburst is also an indicator of imminent visual contact with man’s closest genetic relative: the chimpanzee.

Gombe is the smallest of Tanzania’s national parks: a fragile strip of chimpanzee habitat straddling the steep slopes and river valleys that hem in the sandy northern shore of Lake Tanganyika.

Its chimpanzees – habituated to human visitors – were made famous by the pioneering work of Jane Goodall, who in 1960 founded a behavioural research program that now stands as the longest-running study of its kind in the world. The matriarch Fifi, the last surviving member of the original community, only three-years old when Goodall first set foot in Gombe, is still regularly seen by visitors.

Chimpanzees share about 98% of their genes with humans, and no scientific expertise is required to distinguish between the individual repertoires of pants, hoots and screams that define the celebrities, the powerbrokers, and the supporting characters.

Perhaps you will see a flicker of understanding when you look into a chimp’s eyes, assessing you in return – a look of apparent recognition across the narrowest of species barriers. The most visible of Gombe’s other mammals are also primates. A troop of beachcomber olive baboons, under study since the 1960s, is exceptionallyhabituated, while red-tailed and red colobus monkeys – the latter regularly hunted by chimps – stick to the forest canopy.

The park’s 200-odd bird species range from the iconic fish eagle to the jewel-like Peter’s twinspots that hop tamely around the visitors’ centre.

After dusk, a dazzling night sky is complemented by the lanterns of hundreds of small wooden boats, bobbing on the lake like a sprawling city.

About Gombe Stream National Park

Size: 52 sq km (20 sq miles), Tanzania’s smallest park.
Location: 16 km (10 miles) north of Kigoma on the shore of Lake Tanganyika in western Tanzania.

What to do

Chimpanzee trekking; hiking, swimming and snorkelling;
visit the site of Henry Stanley’s famous “Dr Livingstone I presume” at Ujiji near Kigoma, and watch the renowned dhow builders at work.

When to go
The chimps don’t roam as far in the wet season (February-June, November-mid December) so may be easier to find better picture opportunities in the dry (July-October and late December) to safeguard you and the chimps.

Allow at least 2 days to see them – this is not a zoo so there are no guarantees where they’ll be each day.